![]() ![]() In one astonishing passage, Carroll tells us that if a single E. In thinking about how animal numbers are regulated to avoid over-population, “Elton suggested that, in general, increases in numbers were held in check by predators, pathogens, parasites and food supply.”Įlton’s four regulators are clearly very effective. In his 2016 book The Serengeti Rules, Sean Carroll discusses the work of pioneering ecologist Charles Elton in the 1920s. ![]() The Book of Revelations in the New Testament lists the Four Horsemen of the Apocalypse as conquest, war, famine and death, while in the Old Testament’s Book of Ezekiel they are sword, famine, wild beasts and pestilence or plague.īut whatever we call them, they are remarkably close to what we might call the four horsemen of ecology that regulate population size in nature.
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